Feedback Letter
Response
Of course, such exceptionless laws are fictions. There never was an infinite, flat, frictionless plane on which any object would tend to travel in a straight line until acted on by another force. But laws do tend, in physics, to be more general than in biology. Why is that?
It is because biology covers only a single planet in a vast universe, and each event in biology, such as the evolution of a spine, or adaptation to industrial pollution, is a historically singular event. Generalisations about history are only rules of thumb, and they summarise what is known about past events rather than setting the preconditions for all evolutionary events.
It is true, though, that if the criteria for a natural selection process are fulfilled, the end result has to be natural selection, unless other evolutionary "forces" intervene. But since we cannot predict exactly which mutations will arise, or which alleles will be sampled in a peripheral and isolated population, or what the Sun will do or the tectonic plates will do, in evolutionary theory, we can't make the sorts of predictions we can make in, say, astronomy.
Punctuated Equilibrium (PE) Theory is a generalisation about evolutionary patterns. It says that most change occurs off the fossil record, as it were, because the standard model of evolution of new species has it that speciation occurs in small semi-isolated or completely isolated populations, which are rarely if ever going to be fossilised in the process of speciating, and that species, once adapted to local conditions, will track those conditions rather than adapt much further.
This leads to the "stasis/rapid evolution" pattern in the phylogenetic record. It is not, as such, a "law" about evolution, but a recognition that evolution happens at variable rates, and not over an entire species' range, but locally. So it is a generalisation about (sexual) species evolving over large numbers of species and large periods of time. Unlike a law, a single instance to the contrary is not a falsification of it. Hence, PE is a model of patterns in the evolutionary record, and can be expected to apply only to most speciation events, and not all.
This is true of pretty well all evolutionary rules and models. For that reason, it is not thought that evolution has laws. Ecological theory has laws, though, and they feed into evolutionary theory. So, too, do the laws of physics and chemistry.
That is to say, this is my opinion on the matter.
